Ghost crab digging his hole after the low tide on the white sandy beach, Mahe Seychelles

ghost crab ghost crabs in coastal areas pale nocturnal crab ocypode species decapod crustaceans carapace coloration burrowing habits intertidal zones crablike appearance claw structure quick movements carnivorous diet scuttling behavior remote beaches camouflage ability beach ecosystem complex burrow systems marine invertebrates shell scavengers wavetolerant crabs dune habitat coastal erosion beach conservation breeding season beach pollution effects climate change impacts environmental adaptation beach monitoring nocturnal scavengers resource allocation predatory interactions predation risk shell selectionhigh metabolic rate sand temperature tolerance thermoregulation ghost crab courtship ecosystem engineers coastal ecosystem disturbances population dynamics ghost crab research and conservation digging hole mahe seychelles digging cigarette granite rocks seaweed white sand