Common Mantled Howler resting against branch in forest canopy, close up

alouatta palliata alouatta mantled howler common mantled howler golden mantled howler costa rica wildlife primate monkey biodiversity mammal habitat resting arboreal folivores rainforest canopy jungle national park vocalizations sleeping territorial black howler social behavior protected environment rica guttoral grasping vocal sitting territory ecosystem nature social exotic wild forest howls evergreen growls central sanctuary costa tropical tropics communication america loud