Colourful Chestnut Mandibled Toucan opening large beak and creating sound, animal behavior, close up

ramphastos ambiguus swainsonii toucan costa rica beak yellowthroated central america bird avian frugivorous wildlife perch fauna animal plumage habitat swainson nature vegetation exotic jungle bill environment branch tropical tree large yellow mandibled open looking closeup chestnut bush leaves forest defensive green colorful dark close up black lush mating call behavior creature feather sound creating